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Heyer WR (2005). "Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, Northern South America, and Amazonia". Arquivos de Zoologia (Full text). 37: 269–348. doi:10.11606/issn.2176-7793.v37i3p269-348. Retrieved January 3, 2026.
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