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Weiss, S. J.; Gonçalves, D. V.; Secci-Petretto, G.; Englmaier, G. K.; Gomes-Dos-Santos, A.; Denys, G. P. J.; Persat, H.; Antonov, A.; Hahn, C.; Taylor, E. B.; Froufe, E. (1 March 2021). "Global systematic diversity, range distributions, conservation and taxonomic assessments of graylings (Teleostei: Salmonidae; Thymallus spp.)". Organisms Diversity & Evolution. 21 (1): 25–42. doi:10.1007/s13127-020-00468-7. ISSN 1618-1077. Retrieved 17 October 2025.
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